AmericasAnimaliaArthropodaBeetleCentral AmericaChepo DistrictCocobolo Nature ReserveColeopteraCyclocephala amazonaCyclocephala sp.CyclocephaliniDynastinaeInsectaMYNMaleMasked ChaferPanamaPanama ProvincePolyphagaRhinoceros BeetleScarabaeidaeScarabaeiformiaScarabaeoidea