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Animalia Arthropoda Asia Beetle Cambodia Coleoptera Digitonthophagus gazella Digitonthophagus sp.Insecta Jani Mikkonen Kok Chak Commune MYN Meru Onthophagini Polyphaga Scarabaeidae Scarabaeiformia Scarabaeinae Scarabaeoidea Siem Reap Municipality Siem Reap Province Veal Village

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