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Angkor Angkor Thom Angkor Thom Moat Angkorian Sites Animal Animalia Arthropoda Asia Beetle Biology Cambodia Coleoptera Digitonthophagus gazella Digitonthophagus sp.Imago Insecta MYN Onthophagini Polyphaga Prasat Preah Khan Scarabaeidae Scarabaeiformia Scarabaeinae Scarabaeoidea

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